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The Islamic State has long threatened to strike Russia for helping the Syrian president, Bashar al-Assad, stay in control.

The police presence appeared heavy for the service. Some attendees shouted, “No to war” and “Russia will be free” as they marched to the cemetery where the opposition leader was to be buried.

The reported death of a leading dissident reveals how deeply his crusade against corruption has challenged Russia’s autocratic leader, Vladimir Putin.